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"This is the peer reviewed version of the following article: Heide, T. , Maurer, A. , Eipel, M. , Knoll, K. , Geelvink, M. , Veeck, J. , Knuechel, R. , van Essen, J. , Stoehr, R. , Hartmann, A. , Altmueller, J. , Graham, T. A. and Gaisa, N. T. (2019), Multiregion human bladder cancer sequencing reveals tumour evolution, bladder cancer phenotypes and implications for targeted therapy. J. Pathol., 248: 230-242. doi:10.1002/path.5250 which has been published in final form at https://doi.org/10.1002/path.5250. This article may be used for non-commercial purposes in accordance with Wiley Terms and Conditions for Use of Self-Archived Versions."